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The De Novo Pathway begins with the condensation of L-serine and a fatty-acyl CoA such as palmitoyl-CoA, myristoyl CoA, or stearoyl CoA at the endoplasmic reticulum catalyzed by the enzyme serine palmitoyltransferase (SPT) to form 3-ketosphinganine, which is then reduced to dihydrosphingosine by the NADH-dependent 3-ketosphinganine reductase (KDSR)

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